Portolan Capital Management, LLC

Name: Portolan Capital Management, LLC: Form 13F holdings for the quarter ended 2026-06-30
Creator: Listed

An institutional investment manager in BOSTON, MA. Its Form 13F for the quarter ended Jun 30, 2026, filed Aug 14, 2026. SEC CIK 1350290.

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Reported value$2.67bn
Positions148

Holdings: 148 positions

The filing on sec.gov

As filed, largest first. Reported value is what the manager stated for each position at the quarter end, in US dollars; it is filing data, not a price. Weight is the position's share of the total reported value. Changes compare with the 13F for the quarter ended 2026-03-31.

Portolan Capital Management, LLC, holdings
Issuer	Class	CUSIP	Reported value$m	Weight	Shares	Change on the quarter before
Vishay Precision Group IncVPG	COM	92835K103	151	5.7%	1,008,167	Reduced−11,701
Virtu Finl IncVIRT	CL A	928254101	128	4.8%	2,142,215	Added+930,328
TTM Technologies IncTTMI	COM	87305R109	95.0	3.6%	507,838	Reduced−219,656
Modine MFG CoMOD	COM	607828100	83.8	3.1%	313,683	Reduced−98,409
Qualys IncQLYS	COM	74758T303	72.8	2.7%	529,807	Added+290,779
SHIFT4 PMTS IncFOUR	CL A	82452J109	54.2	2.0%	1,114,116	Added+899,832
Mayville Engr Co IncMEC	COM	578605107	48.7	1.8%	1,299,256	Added+478,120
Enova International IncENVA	COM	29357K103	47.9	1.8%	198,795	Added+117,872
Semtech CorpSMTC	COM	816850101	47.7	1.8%	294,619	Reduced−37,708
Celestica IncCLS	COM	15101Q207	46.3	1.7%	126,856	Reduced−71,007
United Therapeutics Corp DelUTHR	COM	91307C102	44.2	1.7%	81,518	Added+18,083
Genius Sports Limited	SHARES CL A	G3934V109	42.6	1.6%	7,025,809	Added+4,234,791
Powell Inds IncPOWL	COM	739128106	41.7	1.6%	145,631	Added+69,801
Penguin Solutions IncPENG	COM	706915105	41.5	1.6%	546,173	New+546,173
Dave IncDAVE	CLASS A COM NEW	23834J201	39.2	1.5%	105,338	Reduced−104
Flex Ltd	ORD	Y2573F102	37.1	1.4%	229,027	Reduced−214,101
Allient IncALNT	COM	019330109	37.0	1.4%	359,685	Added+36,232
Porch Group IncPRCH	COM	733245104	36.9	1.4%	2,456,451	Added+311,561
Akamai Technologies IncAKAM	COM	00971T101	36.7	1.4%	310,361	Reduced−113,081
Candel Therapeutics IncCADL	COM	137404109	36.0	1.3%	3,490,398	Added+596,073
Talen Energy CorpTLN	COM	87422Q109	34.7	1.3%	90,312	Added+8,738
Regal Rexnord CorporationRRX	COM	758750103	32.8	1.2%	137,498	Added+83,523
Lantheus Holdings IncLNTH	COM	516544103	32.0	1.2%	288,691	Reduced−128,699
Biogen IncBIIB	COM	09062X103	30.9	1.2%	142,856	Added+24,375
Veeco Instrs Inc DelVECO	COM	922417100	30.4	1.1%	401,639	Added+148,260
Luxexperience BVLUXE	SPONSORED ADS	55406W103	30.1	1.1%	4,053,147	Added+319,204
Zeta Global Holdings CorpZETA	CL A	98956A105	29.6	1.1%	1,502,738	Added+1,016,467
Cellebrite Di Ltd	ORDINARY SHARES	M2197Q107	28.2	1.1%	1,932,514	Reduced−248,410
Sportradar Group AG	CLASS A ORD SHS	H8088L103	27.2	1.0%	1,819,949	New+1,819,949
Canada Goose Holdings IncGOOS	SHS SUB VTG	135086106	26.2	1.0%	2,750,523	Reduced−467,754
Bandwidth IncBAND	COM CL A	05988J103	26.1	1.0%	412,701	New+412,701
Warby Parker IncWRBY	CL A COM	93403J106	25.6	1.0%	843,862	Added+343,156
Kontoor Brands IncKTB	COM	50050N103	24.3	0.9%	292,115	New+292,115
Oshkosh CorpOSK	COM	688239201	24.2	0.9%	157,641	Added+139,456
Encore Cap Group IncECPG	COM	292554102	23.9	0.9%	255,715	New+255,715
Twilio IncTWLO	CL A	90138F102	23.8	0.9%	115,397	Added+40,628
Sezzle IncSEZL	COM	78435P105	23.3	0.9%	135,743	Added+65,697
Remitly Global IncRELY	COM	75960P104	22.8	0.9%	1,017,592	Added+319,426
ACM Resh IncACMR	COM CL A	00108J109	22.5	0.8%	177,432	Added+94,176
Element Solutions IncESI	COM	28618M106	22.4	0.8%	469,213	Added+247,981
Nokia CorpNOK	SPONSORED ADR	654902204	21.9	0.8%	1,645,761	Reduced−748,209
Blackberry LtdBB	COM	09228F103	21.7	0.8%	1,714,461	New+1,714,461
International Flavors&FragraIFF	COM	459506101	21.3	0.8%	269,229	New+269,229
Pulse Biosciences IncPLSE	COM	74587B101	20.9	0.8%	753,333	Added+19,961
Moog IncMOG-A	CL A	615394202	20.9	0.8%	49,274	Added+19,837
The Real Brokerage Inc	COM NEW	75585H206	20.1	0.8%	11,042,364	Reduced−3,477,484
Caredx IncCDNA	COM	14167L103	19.5	0.7%	684,805	Added+344,453
PAR Pac Holdings IncPARR	COM NEW	69888T207	19.2	0.7%	341,863	Reduced−351,971
Vishay Intertechnology IncVSH	COM	928298108	18.9	0.7%	351,515	New+351,515
Take-Two Interactive SoftwarTTWO	COM	874054109	18.1	0.7%	72,487	Added+40,268
Gates Indl Corp plc	ORD SHS	G39108108	17.9	0.7%	641,587	New+641,587
Mamas Creations IncMAMA	COM	56146T103	17.6	0.7%	984,869	Reduced−192,067
Kura Sushi USA IncKRUS	CL A COM	501270102	17.4	0.7%	302,010	Added+798
Sanmina CorpSANM	COM	801056102	17.3	0.6%	68,255	Added+40,923
Park Aerospace CorpPKE	COM	70014A104	17.1	0.6%	447,390	Added+36,735
Graham CorpGHM	COM	384556106	16.9	0.6%	136,925	Added+33,369
BEL Fuse IncBELFA	CL B	077347300	15.9	0.6%	47,856	Reduced−5,314
On HLDG AG	NAMEN AKT A	H5919C104	15.6	0.6%	441,137	Added+238,693
Amer Sports Inc	COM SHS	G0260P102	15.3	0.6%	452,880	Added+95,054
Cohu IncCOHU	COM	192576106	15.2	0.6%	206,120	New+206,120
Jabil IncJBL	COM	466313103	15.0	0.6%	38,929	Reduced−38,268
Adeia IncADEA	COM	00676P107	14.6	0.5%	443,060	Reduced−286,401
Vicor CorpVICR	COM	925815102	14.5	0.5%	38,081	Added+11,411
Tenable Holdings IncTENB	COM	88025T102	14.3	0.5%	387,923	New+387,923
GE Healthcare Technologies IGEHC	COMMON STOCK	36266G107	14.2	0.5%	222,393	New+222,393
Crane NXT CoCXT	COM	224441105	13.9	0.5%	271,524	New+271,524
Ranpak Holdings CorpPACK	COM CL A	75321W103	13.9	0.5%	1,894,855	Reduced−52,826
Onto Innovation IncONTO	COM	683344105	13.8	0.5%	36,503	Added+14,240
Genedx Holdings CorpWGS	COM CL A	81663L200	13.8	0.5%	200,315	Added+97,740
Forrester Resh IncFORR	COM	346563109	13.7	0.5%	1,623,925	Added+82,649
Marinemax IncHZO	COM	567908108	12.6	0.5%	344,358	New+344,358
Bio Rad Labs IncBIO	CL A	090572207	12.5	0.5%	42,620	Added+28,447
Quantum CorpQMCO	COM	747906600	11.2	0.4%	1,020,120	New+1,020,120
Baidu IncBIDU	SPON ADR REP A	056752108	11.0	0.4%	95,863	Reduced−270,566
Ferrari N V	COM	N3167Y103	10.8	0.4%	29,079	New+29,079
Digi International IncDGII	COM	253798102	10.6	0.4%	141,243	Reduced−29,013
Nlight IncLASR	COM	65487K100	10.6	0.4%	151,867	Reduced−73,709
Viavi Solutions IncVIAV	COM	925550105	10.5	0.4%	220,118	Added+74,274
Boot Barn Holdings IncBOOT	COM	099406100	10.5	0.4%	63,759	New+63,759
Lightpath Technologies IncLPTH	COM CL A	532257805	10.3	0.4%	629,168	Reduced−411,294
Frequency Electrs IncFEIM	COM	358010106	10.1	0.4%	152,818	Added+2,677
Halozyme Therapeutics IncHALO	COM	40637H109	10.0	0.4%	127,498	New+127,498
Okeanis Eco Tankers Cor	SHS	Y64177101	9.8	0.4%	195,997	Reduced−342,863
Ermenegildo Zegna N V	ORD SHS	N30577105	9.8	0.4%	753,948	Added+43,322
Sweetgreen IncSG	COM CL A	87043Q108	9.1	0.3%	1,029,016	New+1,029,016
Tapestry IncTPR	COM	876030107	9.0	0.3%	61,688	Added+44,650
Solaris Energy Infras IncSEI	COM CL A	83418M103	8.6	0.3%	107,401	Added+79,471
A10 Networks IncATEN	COM	002121101	8.5	0.3%	226,920	New+226,920
Ambarella Inc	SHS	G037AX101	8.3	0.3%	97,072	Reduced−11,677
Tecnoglass Inc	ORD SHS	G87264100	8.3	0.3%	176,254	New+176,254
Quanterix CorpQTRX	COM	74766Q101	8.2	0.3%	1,899,308	Reduced−1,046,515
Tecogen Inc NewTGEN	COM NEW	87876P201	8.1	0.3%	1,551,113	Added+1,107,558
Healthequity IncHQY	COM	42226A107	8.1	0.3%	89,274	Reduced−12,372
Tower Semiconductor Ltd	SHS NEW	M87915274	8.0	0.3%	30,557	Reduced−1,486
Castle Biosciences IncCSTL	COM	14843C105	7.8	0.3%	328,818	Reduced−1,021,465
Silicom Ltd	ORD	M84116108	7.8	0.3%	162,466	New+162,466
Ciena CorpCIEN	COM NEW	171779309	7.5	0.3%	15,272	Reduced−3,194
Ebay Inc.EBAY	COM	278642103	7.3	0.3%	65,741	Reduced−60,355
RB Global IncRBA	COM	74935Q107	7.3	0.3%	63,009	New+63,009
Xometry IncXMTR	CLASS A COM	98423F109	7.1	0.3%	73,163	New+73,163
Mobileye Global IncMBLY	COMMON CLASS A	60741F104	7.0	0.3%	721,338	Added+43,203
Willdan Group IncWLDN	COM	96924N100	6.7	0.2%	84,394	Reduced−9,570
Honest Co IncHNST	COM	438333106	6.4	0.2%	1,740,146	Reduced−1,305,688
Materion CorpMTRN	COM	576690101	6.2	0.2%	20,860	New+20,860
Titan Amer SA	COMMON SHARES	B9151N105	6.0	0.2%	320,509	New+320,509
Global E Online Ltd	SHS	M5216V106	6.0	0.2%	171,521	New+171,521
Meta Platforms IncMETA	CL A	30303M102	5.7	0.2%	10,155	Reduced−106
Celsius Holdings IncCELH	COM NEW	15118V207	5.7	0.2%	194,851	Added+30,254
The Realreal IncREAL	COM	88339P101	5.6	0.2%	478,194	New+478,194
E L F Beauty IncELF	COM	26856L103	5.6	0.2%	75,954	Reduced−60,753
Transmedics Group IncTMDX	COM	89377M109	5.5	0.2%	82,354	Added+71,666
Red Violet IncRDVT	COM	75704L104	5.4	0.2%	84,135	New+84,135
DHT Holdings Inc	SHS NEW	Y2065G121	5.2	0.2%	314,928	Reduced−776,860
Evolv Technologies HLDNGS inEVLV	COM CL A	30049H102	5.2	0.2%	894,478	Reduced−901,210
Procept Biorobotics CorpPRCT	COM	74276L105	5.1	0.2%	227,010	Reduced−24,706
Timken CoTKR	COM	887389104	5.1	0.2%	34,861	New+34,861
Lumentum Holdings IncLITE	COM	55024U109	5.0	0.2%	5,822	Unchanged
Generac Holdings IncGNRC	COM	368736104	4.9	0.2%	16,852	New+16,852
Brunswick CorpBC	COM	117043109	4.8	0.2%	57,175	New+57,175
Equipmentshare Com IncEQPT	COM CL A	29445S100	4.5	0.2%	231,126	New+231,126
Cognyte Software Ltd	ORD SHS	M25133105	4.4	0.2%	500,140	New+500,140
Novanta IncNOVT	COM	67000B104	4.4	0.2%	26,972	Reduced−10,535
Douglas Elliman IncDOUG	COM	25961D105	4.2	0.2%	2,399,364	Reduced−269,666
Bobs Disc Furniture IncBOBS	COM SHS	09681N106	4.0	0.1%	249,906	New+249,906
Everus Constr GroupECG	COM	300426103	3.9	0.1%	23,568	Added+14,843
Methode Electrs IncMEI	COM	591520200	3.8	0.1%	200,000	New+200,000
Fabrinet	SHS	G3323L100	3.6	0.1%	6,418	Reduced−48,345
Ultra Clean Holdings IncUCTT	COM	90385V107	3.5	0.1%	24,831	Reduced−8,869
Kratos Defense & Sec SolutioKTOS	COM NEW	50077B207	3.3	0.1%	65,221	Reduced−194,590
Nextpower IncNXT	CLASS A COM	65290E101	3.2	0.1%	27,101	Reduced−29,646
Hafnia Ltd	SHS	Y2990R101	2.4	<0.1%	367,195	Reduced−1,581,522
Natera IncNTRA	COM	632307104	2.3	<0.1%	8,532	Reduced−2,886
Guardant Health IncGH	COM	40131M109	2.1	<0.1%	13,897	Reduced−258
Gigacloud Technology Inc	CLASS A ORD	G38644103	2.1	<0.1%	65,644	Added+13,630
Alphabet IncGOOGL	CAP STK CL C	02079K107	2.0	<0.1%	5,710	Unchanged
Applied Optoelectronics IncAAOI	COM	03823U102	1.8	<0.1%	11,966	Reduced−17,867
IES Holdings IncIESC	COM	44951W106	1.4	<0.1%	1,940	Reduced−9,929
Casella Waste Sys IncCWST	CL A	147448104	1.4	<0.1%	14,690	Reduced−410
Planet Labs PBCPL	COM CL A	72703X106	1.2	<0.1%	36,869	Reduced−10,419
American Superconductor CorpAMSC	SHS NEW	030111207	1.2	<0.1%	29,314	Reduced−266,322
Dana IncDAN	COM	235825205	0.9	<0.1%	33,121	Reduced−1,116
Digitalbridge Group Inc	CL A NEW	25401T603	0.9	<0.1%	56,763	Reduced−316,367
Lincoln Edl SVCS CorpLINC	COM	533535100	0.7	<0.1%	14,007	Unchanged
L3HARRIS Technologies IncLHX	COM	502431109	0.6	<0.1%	2,157	Unchanged
Open Lending Corp	COM	68373J104	0.6	<0.1%	182,803	Reduced−6,720,085
Telix Pharmaceutical LtdTLX	SPONSORED ADS	87961M105	0.3	<0.1%	25,424	Reduced−515,843
GDS Holdings LtdGDS	SPONSORED ADS	36165L108	0.3	<0.1%	9,120	Reduced−6,161
Talos Energy IncTALO	COM	87484T108	0.2	<0.1%	13,300	Reduced−331,249

Sold out: 32 positions

Held at the end of the quarter before and not reported this quarter.

Portolan Capital Management, LLC, positions sold out
Issuer	Class	CUSIP	Weight	Change on the quarter before
Aerovironment IncAVAV	COM	008073108	0.0%	Sold out−26,880
Amplify Energy Corp NewAMPY	COM	03212B103	0.0%	Sold out−74,120
Amplitude IncAMPL	COM CL A	03213A104	0.0%	Sold out−359,593
Axcelis Technologies IncACLS	COM NEW	054540208	0.0%	Sold out−77,168
Blackline IncBL	COM	09239B109	0.0%	Sold out−159,026
Century Alum CoCENX	COM	156431108	0.0%	Sold out−95,039
Circle Internet Group IncCRCL	COM CL A	172573107	0.0%	Sold out−213,796
Core Natural Resources IncCNR	COM SHS	218937100	0.0%	Sold out−237,036
Crescent Energy CompanyCRGY	CL A COM	44952J104	0.0%	Sold out−2,601,142
Domo IncHUCK	COM CL B	257554105	0.0%	Sold out−197,104
Draganfly Inc.DPRO	COM	26142Q304	0.0%	Sold out−682,730
Establishment Labs Holdings Inc	COM	G31249108	0.0%	Sold out−9,573
Exzeo Group IncXZO	COM SHS	30234F101	0.0%	Sold out−209,885
Forgent Power Solutions IncFPS	COM SHS CL A	34631F102	0.0%	Sold out−117,933
Fulgent Genetics IncFLGT	COM	359664109	0.0%	Sold out−106,642
Green Dot CorpGDOT	CL A	39304D102	0.0%	Sold out−298,697
I3 Verticals IncIIIV	COM CL A	46571Y107	0.0%	Sold out−207,274
Kosmos Energy LtdKOS	COM	500688106	0.0%	Sold out−602,542
Miami International Holdings IncMIAX	COM	59356Q108	0.0%	Sold out−128,819
Murphy USA IncMUSA	COM	626755102	0.0%	Sold out−48,107
Northern Oil & Gas IncNOG	COM	665531307	0.0%	Sold out−204,244
PBF Energy IncPBF	CL A	69318G106	0.0%	Sold out−799,998
Personalis IncPSNL	COM	71535D106	0.0%	Sold out−736,713
Primoris SVCS CorpPRIM	COM	74164F103	0.0%	Sold out−130,626
Propetro HLDG CorpPUMP	COM	74347M108	0.0%	Sold out−314,815
Radnet IncRDNT	COM	750491102	0.0%	Sold out−23,728
Repay Holdings CorpRPAY	COM CL A	76029L100	0.0%	Sold out−2,856,178
Sandridge Energy IncSD	COM NEW	80007P869	0.0%	Sold out−123,591
Sprout Social IncSPT	COM CL A	85209W109	0.0%	Sold out−558,350
Upwork IncUPWK	COM	91688F104	0.0%	Sold out−1,570,560
Vnet Group IncVNET	SPONSORED ADS A	90138A103	0.0%	Sold out−825,490
Xpeng IncXPEV	ADS	98422D105	0.0%	Sold out−275,432

Source: SEC EDGAR, Form 13F-HR, accession 0001104659-26-096980 (on sec.gov). Last ingested by Listed: Oct 22, 2026, 05:26 ET. Listed shows no prices and gives no investment advice.